BA, Biology, Dartmouth College, USA 1975 MD, Stanford University, USA 1982 PhD, Biochemistry, Stanford University, USA 1986
2014 - present Breakthrough Prize in Life Sciences Jury, New York, USA 2016 - present The Rockefeller University, Board of Trustees, New York City, New York, USA 2016 - present Rockefeller Archive Center, Board of Directors, New York City, New York, USA 2016 - present New York Genome Center, Board of Directors, New York City, New York, USA